Papers from the 6th International Conference on Trilobites and their Relatives

Papers from the 6th International Conference on Trilobites and their Relatives By Alan W Owen & David L Bruton 1

Acknowledgements 2

References 2

From Oslo to Prague – the Trilobite meetings 1973–2012 By David L Bruton 5

Introduction 5

Oslo, Norway 1973 (July 1–8) 5

Brock University, St Catharines, Ontario, Canada 1997 (August 22–25) 8

University of Oxford, England, 2001 (April 3–6) 11

Toledo, Spain, 2008 (June 16–24) 14

Prague, Czech Republic, 2012 (July 1–4) 15

Acknowledgements 16

References 16

The 6th International Conference on Trilobites and their Relatives: Tallinn calling, Estonia celebrating By Helje Parnaste 17

Acknowledgements 20

References 21

Early post‐embryonic trilobite stages and possible eggs from the ‘Tu nel Ordovicico del Fabar’ (Middle Ordovician, northwestern Spain) By Enrique Bernardez, Jorge Esteve, Lukaš Laibl, Isabel Rabano and Juan Carlos Gutierrez-Marco 23

Materials and methods 24

Geological setting 24

Trilobite early stages 25

Possible trilobite eggs 27

Discussion 30

Concluding remarks 30

Acknowledgements 31

References 31

Stratigraphy and trilobite biofacies of the Late Ordovician of the Taimyr Peninsula, Arctic Russia By Jan OveR Ebbestad & Richard A Fortey 35

Introduction 35

The Taimyr Peninsula 36

Geography and previous research 36

Structural setting 36

Facies regions and areas 38

Stratigraphy 38

Units, ages and fossils 38

Vesenn’aya Formation (Vesenninskaa tolŝa) 40

Toll Formation (Tollevskaa Svita) 40

Engelgardt Formation (Engel’gardtovskaa Svita) 40

Tolmachev Formation (Tolmač evskaa Svita) 41

Barkov Formation (Barkovskaa Svita) 41

Mutnyj Formation (Mutninskaa svita) 42

Povorotnaya Formation (Povorotninskaa svita) 42

Trilobite distribution and biofacies 43

Stratigraphical distribution 43

Biofacies 43

Scoto‐Appalachian comparison 48

Barkov Formation 49

Conclusions 49

Acknowledgements 49

References 50

The earliest known West Gondwanan trilobites from the Anti‐Atlas of Morocco, with a revision of the Family Bigotinidae Hupe, 1953 By Gerd Geyer 55

Introduction 55

Geological setting and stratigraphy of the Anti‐Atlas sections 56

Tiout section 56

Igoudine Formation 57

Amouslek Formation 57

Tazemmourt section 58

Amouslek section 60

Trilobite occurrence, diversity and biostratigraphy 60

Trilobite preservation and preparation 62

Biostratigraphical implications and biochronological framework 62

Systematic palaeontology 63

Class Trilobita Walch, 1771 64

Order Redlichiida Richter, 1932 64

Suborder Redlichiina Richter, 1932 64

Family Bigotinidae Hupe, 1953 64

Genus Bigotina Cobbold, 1935 65

Bigotina bivallata Cobbold, 1935 73

Bigotina kelleri n. sp 77

Bigotina monningeri n. sp 80

Bigotina sp. A 84

Bigotina sp. B 85

Bigotina angulata Suvorova, 1960 87

Bigotina inornata Egorova, 1986 89

Genus Bigotinella Suvorova, 1960 90

Bigotinella malykanica (Suvorova, 1960 ) 92

Genus Bigotinops Hupe, 1953 92

Bigotinops dangeardi Hupe, 1953 94

Bigotinops chouberti n. sp 97

Genus Ouijjania Hupe, 1953 100

Ouijjania meridionalis Hupe, 1953 102

Genus Demuma Ozdikmen, 2009 104

Demuma nicklesi (Hupe, 1953) 104

‘Demuma bourgini (Hupe, 1953)’ 106

Genus Issendalenia n. gen 106

Issendalenia grandispina n. gen., n. sp 106

Genus Tioutella n. gen 109

Tioutella floccofavosa n. gen., n. sp 109

Genus Pseudobigotina n. gen 111

Pseudobigotina antiatlasensis n. gen., n. sp 113

Genus Hupetina Sdzuy, 1978 115

Hupetina antiqua Sdzuy, 1978 116

Genus Serrania Linan, 1978 119

Serrania verae Linan, 1978 120

Genus Eladiolinania n. gen 120

Eladiolinania castor n. gen., n. sp 122

Eladiolinania pollux n. gen., n. sp 125

Eladiolinania? palaciosi (Linan, Gozalo, Dies Alvarez, Gamez Vintaned & Zamora, 2008) 126

Eladiolinania? gordaensis (Linan, Gozalo, Dies Alvarez, Gamez Vintaned & Zamora, 2008) 127

Genus Suvorovaella n. gen 128

Suvorovaella priva (Suvorova, 1960) 129

Suvorovaella? patria (Suvorova, 1960) 131

Genus and species indeterminate A 131

Genus and species indeterminate B 132

Genus and species indeterminate C 133

Genus and species indeterminate D 133

Genus and species indeterminate E 133

Family Minusinellidae n. fam `134

Minusinella Repina, 1960 134

Family indeterminate 135

Genus and species indeterminate F 135

Superfamily Fallotaspidoidea Hupe, 1953 135

Family Fallotaspididae Hupe 1953 135

Genus Eofallotaspis Sdzuy, 1978 135

Eofallotaspis tioutensis Sdzuy, 1978 137

Eofallotaspis prima Sdzuy, 1978 141

Genus Debrenella n. gen 143

Debrenella larvalis n. gen., n. sp 145

Genus Fallotaspis Hupe, 1953 146

Fallotaspis plana Hupe, 1953, emend 146

Fallotaspis antecedens n. sp 149

Acknowledgements 150

References 150

The nileid trilobite Symphysurus from upper Tremadocian strata of the Moroccan Anti‐Atlas: axonomic reappraisal and palaeoenvironmental implications By Juan Carlos Gutierrez-Marco, Isabel Rabano & Diego C Garcia-Bellido 155

Introduction 155

Materials and methods 156

Stratigraphical context 156

Systematic Palaeontology 157

Order Asaphida Salter, 1864 157

Superfamily ?Cyclopygoidea Raymond, 1925 157

Family Nileidae Angelin, 1854 157

Genus Symphysurus Goldfuss, 1843 157

Symphysurus ebbestadi n. sp 158

Symphysurus sicardi (Bergeron, 1895) 164

Symphysurus? n. sp 166

Notes on palaeoecology and palaeoenvironments 166

Conclusions 168

Acknowledgements 168

References 169

Trilobite‐based biostratigraphy of the Xiaoshiba Lagerstatte By Jin-Bo Hu, Jie Yang, Xi-Guang Zhang, Nigel C Hughes & Tian Lan 173

Introduction 173

Geological setting 174

Results and discussion 176

Systematic palaeontology 178

Class Trilobita Walch, 1771 178

Order Redlichiida Richter, 1933 178

Suborder Redlichiina Harrington, 1959

(in Harrington et al., 1959) 178

Superfamily Ellipsocephalacea Matthew, 1887 178

Family Yunnanocephalidae Hupe, 1953 178

Genus Yunnanocephalus Kobayashi, 1936 178

Yunnanocephalus yunnanensis (Mansuy, 1912) 178

Superfamily Redlichiacea Poulsen, 1927 180

Family Dolerolenidae Kobayashi, 1951 (in Kobayashi & Kato, 1951) 180

Genus Dolerolenus Leanza, 1949 181

Subgenus Dolerolenus (Malungia) Lu, 1961 181

Dolerolenus (Malungia) laevigata Lu, 1961 181

Family Gigantopygidae Harrington, 1959

(in Harrington et al., 1959) 182

Subfamily Yiliangllinae Zhang & Lin, 1980 (in Zhang et al., 1980) 182

Genus Zhangshania Li & Zhang, 1990 (in Li et al., 1990) 182

Zhangshania typica Li & Zhang, 1990 182

Variant of Z. typica 183

Family Metadoxididae Whitehouse, 1939 185

Genus Hongshiyanaspis Zhang & Lin, 1980 185

Hongshiyanaspis yiliangensis Zhang & Lin, 1980 185

Family Redlichiidae Poulsen, 1927 185

Subfamily Wutingaspinae Chang, 1966 185

Genus Chengjiangaspis Zhang & Lin, 1980 185

Chengjiangaspis chengjiangensis Zhang & Lin, 1980 186

Genus Kuanyangia Hupe, 1953 187

Kuanyangia pustulosa Lu, 1941 187

Kuanyangia shaanxiensis Zhang & Lin, 1980 188

Acknowledgements 189

References 189

The taphonomy of a trilobite fauna from an uppermost Katian echinoderm Lagerstatte in South West Wales By Lucy M E McCobb, Patrick D McDermott & Alan W. Owen 193

The Llanddowror bypass echinoderm Lagerstätten 194

Trilobite fauna 194

Taphonomy 195

Conclusions 202

Acknowledgements 202

References 202

The Silurian and Devonian proetid and aulacopleurid trilobites of Japan and their palaeogeographical signifi cance By Christopher P Stocker, Derek J Siveter, Philip D Lane, Mark Williams, Tatsua Ojl, Gengo Tanaka, Toshifumi Komatsu, Simon Wallis, David J Siveter & Th ijs R A Vandenbroucke 205

Introduction 205

Geological background 206

Material and methods 207

Palaeoecology 208

Palaeobiogeography 211

Intra-Japanese terrane palaeobiogeography 211

Connections with Chinese and peri-Gondwanan terranes 211

Systematic palaeontology 212

Order Proetida Fortey & Owens, 1975 212

Family Proetidae Salter, 1864 212

Subfamily Proetinae Salter, 1864 212

Genus Coniproetus Alberti, 1966 212

Coniproetus subovalis (Kobayashi & Hamada,1974) 212

Coniproetus sp A 214

Genus Ganinella Yolkin, 1968 214

Ganinella tenuiceps (Kobayashi & Hamada, 1987) 215

Ganinella fukujiensis (Kobayashi & Hamada,1977) 216

Ganinella oisensis (Kobayashi & Hamada, 1977) 218

Genus Gomiites Pr˘ibyl & Vanĕk, 1978 219

Gomiites granulatus (Kobayashi & Hamada, 1974) 220

Gomiites latiaxis (Kobayashi & Hamada, 1986) 222

Subfamily Crassiproetinae Osmolska, 1970 223

Genus Hedstroemia Pr˘ibyl & Vanĕk, 1978 223

Hedstroemia sugiharensis (Kobayashi & Hamada,1974) 223

Subfamily Cornuproetinae Richter et al in Moore, 1959 224

Genus Interproetus Šnajdr, 1977 224

Interproetus mizobuchii n. sp 224

Subfamily Eremiproetinae Alberti, 1967 226

Genus Eremiproetus Richter & Richter, 1919 226

Eremiproetus? subcarinatus (Kobayashi & Hamada, 1974) 226

Eremiproetus? magnicerviculus (Kobayashi & Hamada, 1974) 227

Subfamily Dechenellinae Pr˘ibyl, 1946 227

Genus Dechenella Kayser, 1880 227

Dechenella minima Okubo, 1951 227

Subfamily Warburgellinae Owens, 1973 228

Genus Latiproetus Lu, 1962 228

Latiproetus bilobus (Kobayashi & Hamada, 1974) 228

Order Aulacopleurida Adrain, 2011 229

Family Aulacopleuridae Angelin, 1854 229

Subfamily Otarioninae Richter & Richter, 1926 229

Genus Maurotarion Alberti, 1969 229

Maurotarion megalops (Kobayashi & Hamada, 1977) 229

Acknowledgements 230

References 230