Mesozoic Fishes 5 – Global Diversity and Evolution

ADAMS, Colin E., Scottish Centre for Ecology & the Natural Environment, University of Glasgow, Glasgow, Scotland. ALVARADO-ORTEGA, Jesús, Instituto de Geologia, Universidad Nacional Autónoma de México; Ciudad Universitaria, Delegación, Coyoacán, México, México. AMARAL, Cesar R. L., Instituto de Biologia Roberto Alcântara Gomes, Universidade do Estado do Rio de Janeiro, São Francisco Xavier, Maracanã, RJ, Brazil. ARRATIA, Gloria, Natural History Museum and Biodiversity Institute, The University of Kansas, Dyche Hall, Lawrence, Kansas, U.S.A. BERRELL, Rodney W., Macquarie University, Sydney, Australia. BORDEN, W. Calvin, Department of Biology, Loyola University Chicago, Chicago, Illinois U.S.A. BRINKMAN, Donald B., Royal Tyrrell Museum of Palaeontology, Drumheller, Alberta, Canada. BRITO, Paulo M., Instituto de Biologia Roberto Alcântara Gomes, Universidade do Estado do Rio de Janeiro, São Francisco Xavier, Maracanã, RJ, Brazil. CHALLANDS, Thomas James, School of Geosciences, Grant Institute of Earth Sciences, University of Edinburgh, Edinburgh, Scotland. CHATTERTON, Brian D. E., Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, Canada. CIONE, Alberto Luis, División Paleontología Vertebrados, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata, Argentina. CUMBAA, Stephen L., Palaeobiology, Canadian Museum of Nature, Ottawa, Ontario, Canada. DAVIS, Matthew P., The Field Museum, Department of Zoology, Chicago, Illinois, USA. ESPINOSA-ARRUBARRENA, Luis, Museo de Geología. Universidad Nacional Autónoma de Mexico. FIELITZ, Christopher, Emory & Henry College, Emory, Virginia, U.S.A. FOWLER, Denver W., Museum of the Rockies, and Department of Earth Sciences, Montana State University, Bozeman, U.S.A. FREEDMAN, Elizabeth A., Museum of the Rockies, and Department of Earth Sciences, Montana State University, Bozeman, U.S.A. GIBB, Stacey, Department of Earth and Atmospheric Sciences, University of Alberta, Edmonton, Canada. GONZÁLEZ-BARBA, Gerardo, Museo de Historia Natural, Universidad Autónoma de Baja California Sur, La Paz, B.C.S., Mexico. GONZÁLEZ-RODRÍGUEZ, Katia A., Instituto de Ciencias Básicas e Ingeniería, Museo de Paleontología, Centro de Investigaciones Biológicas, Universidad Autónoma del Estado de Hidalgo, Mineral de la Reforma, Hidalgo, Mexico. GOUIRIC-CAVALLI, Soledad, División Paleontología Vertebrados, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, La Plata, Argentina. GRANDE, Terry, Department of Biology, Loyola University Chicago, Chicago, Illinois, U.S.A. IRVIN, Kelly J., Arkansas Game & Fish Commission, Benton, Arkansas, U.S.A. KAISER, Thomas, Zoological Institute and Museum, University of Hamburg, Hamburg, Germany. KEMP, Anne, Australian Rivers Institute, Nathan Campus, Griffith University, Brisbane, Queensland, Australia. LISTON, Jeff, Department of Natural Sciences, National Museums Scotland, Edinburgh, Scotland; School of Earth Sciences, Wills Memorial Building, Queens Road, University of Bristol, England; Division of Environmental & Evolutionary Biology, School of Life Sciences, College of Medical Veterinary & Life Sciences, University of Glasgow, Glasgow, Scotland. MARTÍN-ABAD, Hugo, Unidad de Paleontología, Departamento de Biología, Facultad de Ciencias, Universidad Autónoma de Madrid, Cantoblanco, Madrid, Spain. MICKLE, Kathryn, University of Kansas, Biodiversity Research Institute, Lawrence, Kansas, U.S.A.; Rockhurst University, Biology Department, St. Ignatius Science Building, Kansas City, Missouri, U.S.A. MURRAY, Alison M., Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada. NEUMAN, Andrew G., Royal Tyrrell Museum of Palaeontology, Drumheller, Alberta, Canada. NEWBREY, Michael G., Royal Tyrrell Museum of Palaeontology, Drumheller, Alberta, Canada; Department of Biological Sciences, University of Alberta, Edmonton, Alberta, Canada. POYATO-ARIZA, Francisco José, Unidad de Paleontología, Departamento de Biología, Facultad de Ciencias, Universidad Autónoma de Madrid, Cantoblanco, Madrid, Spain. SCHRÖDER-ADAMS, Claudia J., Department of Earth Sciences, Carleton University, Ottawa, Ontario, Canada. SCHULTZE, Hans-Peter, Natural History Museum and Biodiversity Institute, The University of Kansas, Dyche Hall, Lawrence, Kansas, U.S.A. SMITH, W. Leo, Division of Fishes, Field Museum of Natural History, Chicago, Illinois, U.S.A. TISCHLINGER, Helmut, Stammham, Germany. UNDERWOOD, Charlie J., School of Earth Sciences, Birkbeck College, London, UK. WILSON, Mark V. H., Department of Biological Sciences, University of Alberta, Edmonton, Canada. WINKLER, Dale A., Department of Earth Sciences, Southern Methodist University, Dallas, Texas, U.S.A. WOODWARD, Holly N., Museum of the Rockies, and Department of Earth Sciences, Montana State University, Bozeman, U.S.A. YABUMOTO, Yoshitaka, Department of Natural History, Kitakyushu Museum of Natural History and Human History, Kitakyushu, Fukuoka, Japan.

Preface 7
Acknowledgements 8
Katia A. GONZÁLEZ-RODRÍGUEZ, Luis ESPINOSA-ARRUBARRENA and Gerardo GONZÁLEZ-BARBA
An overview of the Mexican fossil fish record 9–34
Kathryn E. MICKLE
Revisiting the actinopterygian preoperculum 35–71
PDF Hugo MARTÍN-ABAD and Francisco José POYATO-ARIZA
Amiiforms from the Iberian Peninsula: historic review and research prospects 73–86
PDF Gloria ARRATIA and Hans-Peter SCHULTZE
Outstanding features of a new Late Jurassic pachycormiform fish from the Kimmeridgian of Brunn, Germany and comments on current understanding of pachycormiforms 87–120
Jeff LISTON
The plasticity of gill raker characteristics in suspension feeders: Implications for Pachycormiformes 121–143
Jeff LISTON, Michael G. NEWBREY, Thomas James CHALLANDS and Colin E. ADAMS
Growth, age and size of the Jurassic pachycormid Leedsichthys problematicus Osteichthyes: Actinopterygii) 145–175
Soledad GOUIRIC-CAVALLI and Alberto Luis CIONE
“Pholidophorus argentinus” DOLGOPOL DE SAEZ, 1939 from Upper Jurassic beds of the Neuquén Province of Argentina is not a pholidophoriform but an aspidorhynchid (Actinopterygii, Aspidorhynchiformes) 177–186
Hans-Peter SCHULTZE and Gloria ARRATIA
The caudal skeleton of basal teleosts, its conventions, and some of its major evolutionary novelties in a temporal dimension 187–246
Kelly J. IRWIN and Christopher FIELITZ
Ichthyodectiform fishes from the Late Cretaceous (Campanian) of Arkansas, USA 247–266
Alison M. MURRAY and Mark V. H. WILSON
Two new paraclupeid fishes (Clupeomorpha: Ellimmichthyiformes) from the Upper Cretaceous of Morocco 267–290
Michael G. NEWBREY Donald B. BRINKMAN, Dale A. WINKLER, Elizabeth A. FREEDMAN, Andrew G. NEUMAN, Denver W. FOWLER and Holly N. WOODWARD
Teleost centrum and jaw elements from the Upper Cretaceous Nemegt Formation (Campanian–Maastrichtian) of Mongolia and a re-identification of the fish centrum found with the theropod Raptorex kreigsteini 291–303
Cesar R. L. AMARAL, Jesús ALVARADO-ORTEGA and Paulo M. BRITO
Sapperichthys gen. nov., a new gonorynchid from the Cenomanian of Chiapas, Mexico 305–323
Matthew P. DAVIS, Gloria ARRATIA and Thomas M. KAISER
The first fossil shellear and its implications for the evolution and divergence of the Kneriidae (Teleostei: Gonorynchiformes) 325–362
Michael G. NEWBREY, Alison M. MURRAY, Mark V. H. WILSON, Donald B. BRINKMAN and Andrew G. NEUMAN
A new species of the paracanthopterygian Xenyllion (Sphenocephaliformes) from the Mowry Formation (Cenomanian) of Utah, USA 363–384
Terry GRANDE, W. Calvin BORDEN and W. Leo SMITH
Limits and relationships of Paracanthopterygii: A molecular framework for evaluating past morphological hypotheses 385–418
W. Calvin BORDEN, Terry GRANDE and W. Leo SMITH
Comparative osteology and myology of the caudal fin in the Paracanthopterygii (Teleostei: Acanthomorpha) 419–455
Katia A. GONZÁLEZ-RODRÍGUEZ, Hans-Peter SCHULTZE and Gloria ARRATIA
Miniature armored acanthomorph teleosts from the Albian/Cenomanian (Cretaceous) of Mexico
Yoshitaka YABUMOTO and Paulo M. BRITO
The second record of a mawsoniid coelacanth from the Lower Cretaceous Crato Formation, Araripe Basin, northeastern Brazil, with comments on the development of coelacanths 489–498
Anne KEMP and Rodney W. BERRELL
Lungfish as environmental indicators 499–508
Stephen L. CUMBAA, Charlie J. UNDERWOOD and Claudia J. SCHRÖDER-ADAMS
Paleoenvironments and Paleoecology of the Vertebrate Fauna from a Late Cretaceous Marine Bonebed, Canada 509–524
Alison M. MURRAY, Mark V. H. WILSON, Stacey GIBB and Brian D. E. CHATTERTON
Additions to the Late Cretaceous (Cenomanian/Turonian) actinopterygian fauna from the Agoult locality, Akrabou Formation, Morocco, and comments on the palaeoenvironment 525–548
Helmut TISCHLINGER and Gloria ARRATIA
Ultraviolet light as a tool for investigating Mesozoic fishes, with a focus on the ichthyofauna of the Solnhofen archipelago 549–560

Mesozoic Fishes meetings are designed to bring together researchers, students, and other interested persons every four years to present research and to foster discussion and collaboration about fossil fishes of the Mesozoic. The meetings often also include presentations on related subjects such as early Cenozoic fishes. The first four meetings were all held in Europe (Eichstätt, Germany, 1993; Buckow (near Berlin), Germany, 1997; Serpiano (near Lugano), Switzerland, 2001; Miraflores de la Sierra (near Madrid), Spain, 2005). Therefore, it was a bold and exciting departure for our Mexican hosts to invite us all to Saltillo, Coahuila state, Mexico, for the 5th Mesozoic Fishes meeting, which was originally planned for August/September 2009. This would be the first Mesozoic Fishes meeting to be held outside of Europe. The year 2009 proved to be a difficult one for the organization of the meeting for reasons beyond their control. One issue might have been news reports about drug-related violence in northern Mexico, leading potential attendees to be nervous about travel to the area. However, a bigger problem was an outbreak of so-called “swine flu” or “swine influenza” which was initially recognized in Mexico in April 2009 and, despite its name, it was spread from human to human, and later became a pandemic. Government health agencies recommended against travel to Mexico that spring and summer. In the end, there were too few registrants for the meeting to be held in 2009. The organizers took the difficult decision to cancel the meeting for 2009 and to organize it instead for the summer of 2010. This decision involved many difficulties including re-booking of hotels, meeting venues, social events, and excursions, and re-applying for financial and logistical support from a variety of Mexican government and academic institutions. Fortunately for the Mesozoic Fishes community, and especially for those of us who attended, the organizers were able to accomplish all of the re-scheduling, and to invite us to Mesozoic Fishes 5 in Saltillo during August 2-7, 2010. By that time, conditions were much better for travel, and the number of registrants was larger than the previous year. The meetings in Saltillo were held at the Museo del Desierto, a modern and beautiful facility with excellent exhibits about the fauna and flora of the region and featuring attractive gardens and beautiful views of the city and the surrounding valley and mountains. The official hosts included the Secretaría de Educación y Cultura del Estado de Coahuila, the Museo del Desierto itself in Saltillo, the Universidad Nacional Autónoma de México – Instituto de Geología, and the Museo de Paleontología of the Universidad Autónoma del Estado de Hidalgo. The organizing committee included María del Rosario Gómez-Núñez, Katia A. González-Rodríguez, and Jesús Alvarado-Ortega. One of the most exciting things about the conference was meeting the large number of Mexican colleagues and students, and learning about their work and the very interesting fossil sites that are being studied in that country. The proportion of young scientists from Mexico and from elsewhere in Latin America was much higher than we have seen elsewhere, a fact that bodes well for the future of paleoichthyology in Central and South America. Mexican colleagues dedicated the meeting to the memory of the late Shelton P. Applegate, who stimulated research, collections, and graduate education based on Mexican fossil fishes over several decades. Shelley attended previous Mesozoic Fishes meetings, and died in 2005. Presentations at the meeting were organized into five days of oral and poster sessions concerning fish taxa ranging taxonomically from chondrichthyans to teleosts and fossil assemblages ranging in age from Triassic to Cenozoic. Many important discoveries were announced and discussed, new research techniques were applied, and synthetic reviews of fossil sites were presented. The present volume is based largely on work that was presented at Mesozoic Fishes 5 in Saltillo, with additional contributions relevant to the theme of Mesozoic Fishes. Special events included the opening of a new exhibition of Mexican fossil fishes at the Museo del Desierto; many of the fishes in the exhibition were the subject of presentations during the meeting. A memorable group dinner where we were entertained by a Mariachi band was held in a nearby restaurant. Excursions included a city tour with a visit to the central marketplace and a visit to the “Museum of the Dead” where we learned the meaning of being “like water for chocolate”, which means being exactly or perfectly ready. The field trip on the last day to Late Cretaceous sites in Coahuila state included visits to local museums and localities, a walk in the desert at Porvenir de Jalpa to view dinosaur trackways, a chance to see petroglyphs at Narihua, and a farewell banquet hosted by the people of the town General Cepeda that included musical and dance performances, followed by general dancing. The meeting as a whole was very well organized, and participants felt warmly welcomed by the their Mexican hosts. We left Saltillo with a new appreciation of the exciting work being done in Mexico, with much to think about scientifically, with new opportunities for collaboration, and with new friends and renewed old friendships.